New barklice (Psocodea, Trogiomorpha) from Lower Cretaceous Spanish amber

Barklice are insects belonging to the order Psocodea. They herbivorous or detritivorous, and inhabit a wide range of environments. Their oldest fossil record dates back late Carboniferous, but it was not until Cretaceous that they became much more diverse. However, their could be affected by taphonomic processes due tiny size soft bodies. Here, we present new psocid specimens from five amber-bearing outcrops in Spain Albian age. One specimens, well-preserved nymph assigned †Archaeatropidae, lacks evidence debris-carrying behaviour. Some belong previously known species Archaeatropos alavensis Baz & Ortuño Preempheria antiqua (Trogiomorpha: Atropetae), providing anatomical biogeographical information. Furthermore, describe species, Libanoglaris hespericus sp. nov. (†Archaeatropidae). The diagnosis for family †Archaeatropidae is emended. abundance psocids amber virtual absence compression bias. Considerations on phylogenetic placement trogiomorphan families relationships between †Empheriidae included. Today, least diverse suborder Trogiomorpha, this comprises majority psocodean described date, possibly palaeobiological evolutionary constraints. Trogiomorphans have been relegated marginal habitats niche competition with psocomorphans. Debris-carrying behaviour response predatory pressure might widely distributed, geographically phylogenetically, nymphs.